The Division of Environmental Biology (DEB) Coresupports research and training on evolutionary and ecological processes acting at the level of populations, species, communities, and ecosystems. DEB encourages research that elucidates fundamental principles that identify and explain the unity and diversity of life and its interactions with the environment over space and time. Research may incorporate field, laboratory, or collection-based approaches; observational or manipulative studies; synthesis activities; phylogenetic discovery projects; or theoretical approaches involving analytical, statistical, or computational modeling. Proposals should be submitted to the core clusters (Ecosystem Science, Evolutionary Processes, Population and Community Ecology, and Systematics and Biodiversity Science). DEB also encourages interdisciplinary proposals that cross conceptual boundaries and integrate over levels of biological organization or across multiple spatial and temporal scales.Research addressing ecology and ecosystem science in the marine biome should be directed to the Biological Oceanography Program in the Division of Ocean Sciences; research addressing evolution and systematics in the marine biome should be directed to the Evolutionary Processes or Systematics and Biodiversity Science programs in DEB. All DEB programs also encourage proposals that leverage NSF-supported data networks, databases, centers, and other forms of scientific infrastructure, including but not limited to the National Ecological Observatory Network (NEON), Long-Term Ecological Research (LTER), Environmental Data Initiative (EDI), and Integrated Digitized Biocollections (iDigBio). The Division of Environmental Biology seeks to strengthen the U.S. Environmental Biology workforce by increasing the participation of all individuals in science. DEB is dedicated to expanding traditional broader impacts and supporting proposals that include inclusive and effective efforts to recruit and retain biology students, postdoctoral researchers, and early-investigators from groups historically underrepresented in the biological sciences.